1980-1981: “A” level degree in Biology, Chemistry and Mathematics, Moraitis School, Athens, Greece.
1984: Bachelor of Science (B.Sc.) in Cell Biology with Honours Class I, University of Essex, Essex, U. K.
1985:
Diplome d’ Etudes Approfondies (D.E.A.) in Microbiology, Institut de Microbiologie, Centre d' Orsay, Université de Paris-XI, Orsay, France.
1989:
Thèse de Doctorat (Ph.D) in “Science de la vie” (Molecular Biology)
with mention “Trés Honorable”, Université de Paris XI, Orsay, France.